Image Credit: Mecum.Some muscle cars are rare because relatively few were built. Others become genuinely special when the right engine, transmission, options, colors, and surviving components all come together, and this 1969 Plymouth Road Runner firmly belongs in the latter category.At first glance, it's a wonderfully understated example. There's black paint, a black interior, simple steel wheels with chrome hubcaps, and none of the visual excess you might expect from something packing one of Chrysler's most legendary engines.Under the Air Grabber hood sits the reason collectors will pay close attention when this Plymouth crosses the auction block. It's powered by a matching-numbers 426 Hemi V8, which is connected to the original four-speed manual transmission it left the factory with.AdvertisementAdvertisementThe car is scheduled to go under the hammer at Mecum's Nashville auction on September 26. No estimate has been published, but given the specification and rarity involved, this isn't likely to be one of the cheaper Road Runners changing hands that weekend.The 426 Hemi Makes All The DifferenceImage Credit: Mecum.Plymouth introduced the Road Runner for 1968 as a relatively affordable muscle car, and buyers quickly embraced the formula. Sales climbed to roughly 81,000 examples for 1969, making it the model's most successful year.Most weren't Hemis. The vast majority left the factory with the standard 383-cubic-inch V8, while the expensive 426 Hemi sat at the top of the performance hierarchy with a factory rating of 425 horsepower.According to production figures cited for 1969, only 787 Road Runners received the Hemi across the available body styles. That already makes any genuine Hemi Road Runner an uncommon machine before transmission, equipment, and configuration enter the equation.AdvertisementAdvertisementMecum lists this particular car as one of just 234 examples produced with a four-speed manual transmission. It also retains its matching-numbers 426 V8 and matching-numbers A833 four-speed, making its mechanical pedigree a major part of the appeal.Super Track Pak Adds Some Serious HardwareImage Credit: Mecum.The original buyer didn't stop after checking the box for the Hemi. This Road Runner was also ordered with the A34 Super Track Pak, giving it hardware intended to make better use of all that V8 muscle.That package includes a Dana 60 rear axle with aggressive 4.10 gearing, along with other heavy-duty equipment. The car also has a 26-inch radiator and power front disc brakes, while Mecum says its suspension has been rebuilt.Naturally, there's a Hurst shifter for the four-speed. Firestone Redline tires wrap around black 15-inch steel wheels, preserving the stripped-down appearance that made the Road Runner such an appealing counterpoint to increasingly luxurious muscle cars.AdvertisementAdvertisementPeriod details include an AM radio, three-speed windshield wipers, and, of course, the Road Runner's famous "Beep Beep" horn.Matching Numbers And Original DetailsImage Credit: Mecum.This example underwent a restoration in 2015, but several important original identifiers remain. Mecum notes that it retains its original fender tag, matching original trunk drip-rail stamp, and matching original radiator-core stamp.The odometer currently displays 32,415 miles, although Mecum explicitly notes that this figure only represents what is shown and shouldn't be interpreted as verified mileage.A Dave Wise vehicle inspection report accompanies the car as well. Combined with the matching drivetrain and surviving factory markings, that should give serious bidders plenty to examine before auction day.A Serious Mopar CollectibleA black-on-black Hemi Road Runner with a four-speed and Super Track Pak checks an impressive number of boxes for Mopar collectors. Its relatively restrained appearance arguably makes the specification even better, since little beyond the hood tells casual observers what's lurking underneath.AdvertisementAdvertisementMecum hasn't published a pre-auction estimate, so predicting the final number would be speculation. Six figures would hardly be surprising, though, considering genuine Hemi-powered Mopars occupy a very different corner of the collector market from ordinary Road Runners.For anyone hunting a serious late-1960s Mopar, September 26 could get expensive very quickly.If you want more stories like this, follow Guessing Headlights on Yahoo so you don't miss what's coming next.
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